The name "hornwort" refers to the long thin
sporophytes which are embedded in the top of the plant and remain attached to it and continue to grow throughout its life. As all
bryophytes, hornworts are
gametophyte dominant.
Other things which make the hornwort different when viewed under a microscope, are that the spaces between the cells are filled with
mucilage rather then air chambers, where the
cyanobacteria of the genus Nostoc live. The cyanobacteria fix nitrogen, which is used by the plant, in return for
carbohydrates.
They also have a single, large
chloroplast in each cell often completely enclosing the cell
nucleus, which is unique among plants. This structure imparts a particular quality of colour and translucency to the body (
thallus) of the plant. One chloroplast per cell is common in many
algae.
The exact age of hornworts is not known, due to few fossil records having been found, but are believed to have evolved around 380 million years ago. There are about 100 species of hornworts, found worldwide in temperate and tropical regions on tree trunks, riverbanks, and other damp locations.
Cyanobacteria
Hornworts are similar in appearance to the thallose liverworts and are difficult to recognise if the sporophytes are not present their gametophyte range in size from 10 mm to 20 mm along their longest axis. Hornworts are green and strap shaped and do not have any leaves or stem but are attached to the ground by
rhizoids. The Sporophytes are long, horn-shaped, and green with brown tips when releasing
spores.